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Molewaterplein 40, 3015 GD Rotterdam, The Netherlands Tel: +31-107032861, Fax: +31-107033639, E-mail: a.vanderlelij@erasmusmcnl Received: October 30, 2023 Revised: November 23, 2023 Accepted: December 6, 2023 Copyright 2024 Korean Endocrine Society This is an Open Access article distributed under the terms of the Creative Commons Attribution Non-Commercial License ( which permits unrestricted non-commercial use, distribution, and reproduction in any medium, provided the original work is properly cited

Histones including H1, H2A, H2B, H3 and H4 are known positively charged proteins which can bind tightly to the negatively charged DNA, favoring histone-DNA complex formation (also named nucleosome) [61]